Tetanurae Gauthier, 1986
Definition- (Passer domesticus <- Ceratosaurus nasicornis)
(Holtz et al., 2004; modified from Padian et al., 1999; modified from Gauthier,
1986)
Other definitions- (Allosaurus fragilis + Passer domesticus)
(modified from Novas, 1992)
(Passer domesticus <- Torvosaurus tanneri) (modified from Sereno,
1998)
(Passer domesticus <- Ceratosaurus nasicornis, Carnotaurus
sastrei) (Sereno, in press)
= Intertheropoda Paul, 1988
= Avipoda Novas, 1992
= Tetanurae sensu Sereno, in press
Definition- (Passer domesticus <- Ceratosaurus nasicornis,
Carnotaurus sastrei)
Comments- The addition of Carnotaurus as an external specifier
by Sereno (in press) seems counter-productive. Tetanurae was designed as a stem
away from Ceratosaurus, and abelisaurids were not explicitly discussed
(having been named only a year prior). In fact, technically, Indosaurus
and Indosuchus were classified as tetanurines by Gauthier (1986), since
he lists them as carnosaurs. If abelisaurids are megalosauroids/torvosauroids
(as in Paul, 1988), it shouldn't stop megalosauroids/torvosauroids from being
tetanurines.
Dandakosaurus Yadagiri, 1982
D. indicus Yadagiri, 1982
Early Jurassic
Kota Formation, India
Holotype- (GSI 1/54Y/76) proximal pubis
Paratypes- ?(GSI coll.) teeth, dorsal vertebra (160 mm), proximal caudal
vertebra (150 mm), proximal ischium
Diagnosis- open obturator notch in pubis; anteroproximal flaring and
smoothly convex proximal surface of pubis; more proximally placed obturator
notch than Patagonykus.
Description- The illustrated tooth is typical of most theropods in being
laterally compressed, recurved and having fine serrations. The tooth seems more
compressed (30% of FABL) than most theropods (eg. Liliensternus airelensis,
Gojirasaurus, Dilophosaurus, Magnosaurus, Torvosaurus, Szechuanosaurus),
though Liliensternus liliensterni is similar in this regard. Though interdental
variation could be a factor here, as posterior teeth are known to be more laterally
compressed in theropods, Dilophosaurus and Magnosaurus never reach
a Dandakosaurus level of compression anywhere in the tooth row.
The dorsal vertebra is opisthocoelous and lacks a pleurocoel. Opisthocoelous
dorsals are only known in non-coelurosaurian tetanurines (the exception is Mononykinae),
suggesting Dandakosaurus is a member of this clade. The absence of a
pleurocoel is of little use without positional data.
The caudal vertebra is amphicoelous, with "two lateral cavities on either
side" and a keeled ventral surface. The first character is common in theropods.
The second is only known in Carcharodontosaurus, Acrocanthosaurus, Patagonykus,
Spinostropheus, Nomingia, caenagnathoids and Achillobator. Few theropods
are reported to have ventral keels on their proximal caudals, including "Capitalsaurus",
Sinraptor dongi, Bagaraatan, Inosaurus, Spinostropheus, alvarezsaurids
and SQU-2-7, an isolated caudal from the Cretaceous of Oman.
The holotype proximal pubis has several odd characters. There is no obturator
fenestra, just an open obturator notch, as in Elaphrosaurus, Eustreptospondylus,
Suchomimus, allosaurids, carcharodontosaurids and coelurosaurs. The proximal
border forms a smooth convex arch from the ilial contact to the ischia contact,
with no distinct peduncles. This is approached in some coelophysoids (eg. Coelophysis
longicollis referred specimen) and Archaeornithomimus, but is most
similar to Patagonykus, Unenlagia and Achillobator. Proximally,
the pubis flares sharply anteriorly to form an acute anteroproximal corner.
This is similar to the situation in Coelurus, tyrannosaurids, Caudipteryx,
Nanshiungosaurus and alvarezsaurids. It seems to only be developed in forms
with somewhat mesopubic pelvic orientations. Additional evidence for mesopuby
in Dandakosaurus may come from the possible pubic peduncle of the ischium,
which fits the ischial surface of the pubis to form an angle of about 35 degrees
between the bones.
Only the proximal portion of the ischium is preserved. There is a concave anterior
margin which matches up with the ischial peduncular area of the pubis. No other
area is appropriate for the pubic contact. If the concave area were the acetabulum,
the pubis and ischium would be subparallel. Articulating the pubis more dorsoposteriorly
on the ischium leaves no acetabular area leaves nowhere for an acetabular surface
or ischial peduncle. The acetabular surface is small and sharply concave, so
may be
broken and continuous with the ischial peduncle in life. The posterior surface
is gently concave, while the anterior surface is broken.
Comments- The phylogenetic relationships of this taxon are uncertain,
especially considering how advanced it seems for its age. However, I have heard
that the Kota Formation could extend into the Middle or Late Jurassic. It seems
to be a tetanurine based on the opisthocoelous dorsal centra. The only coelurosaurs
with such centra are mononykines, which have procoelous caudal centra, though
they have ventrally keeled caudals, an open pubic obturator notch, and an anteroproximally
flared pubis. In addition, Patagonykus shares the presence of lateral
fossae in the caudal centra and a smoothly convex proximal pubic border. Unfortunately,
Patagonykus and Alvarezsaurus show the procoelous caudals evolved
before the opisthocoelous dorsals in alvarezsaurids. Also, the dental morphology
is less derived than one would expect for even a basal alvarezsaurid. Thus,
Dandakosaurus is unlikely to be alvarezsaurid. The caudal fossae, open
pubic obturator notch, and mesopubic pelvis are similar to Achillobator,
but deinonychosaurs lack opisthocoelous dorsals and have thicker teeth. Thus,
Dandakosaurus represents a mesopubic tetanurine of uncertain affinities,
probably not coelurosaurian. It is one of the earliest tetanurines, along with
"Saltriosaurus" and "Merosaurus". Previous suggestions regarding
abelisauroid affinity (Aravind, DML 1997) were based on maxillary characters
unknown in the specimen. Where Aravind got his data is unknown, but it can be
considered irrelevant now that the known material has been determined.
Reference- Yadagiri, 1982. Osteological studies of a carnosaurian dinosaur
from the Lower Jurassic Kota Formation: Andhra Pradesh. Geological Survey of
India (Progress Report for Field Season Programme 1981-1982), Regional Palaeontological
Laboratories, Southern Region. 7 pp.
http://dml.cmnh.org/1997May/msg00840.html
Duriavenator Benson, 2008
= "Walkersaurus" Welles, Powell and Pickering vide Pickering, 1995
D. hesperis (Waldman, 1974) Benson, 2008
= Megalosaurus hesperis Waldman, 1974
= "Walkersaurus" hesperis (Waldman, 1974) Welles, Powell and
Pickering vide Pickering, 1995
Late Bajocian, Middle Jurassic
Upper Inferior Oolite, England
Holotype- (BMNH R332) (~5 m) partial premaxillae, maxilla, vomer, partial
dentaries, partial surangular, teeth, fragments
Diagnosis- (after Benson, 2008) deep groove on dorsal surface of jugal
process containing numerous pneumatic foramina; array of small foramina in ventral
part of articular surface for premaxilla.
Comments- This specimen was originally described by Owen (1883) and referred
to Megalosaurus bucklandi. Walker (1964) noted it was probably a distinct
species, due to tooth count and a supposed lateral groove between the premaxilla
and maxilla. Waldman (1974) officially named the species Megalosaurus hesperis.
Pickering (1995) listed "Walkersaurus" hesperis, attributed
to Welles, Powell and Pickering. He later (Welles and Pickering, 1999) mentioned
the taxa in his description of Dilophosaurus "breedorum" and
possibly in his identically titled redescription of Megalosaurus. However,
none of Pickering's publications are valid according to the ICZN, as they violate
Article 8.1 and Recommendation 8A. Specifically, it was not "produced in
an edition containing simultaneously obtainable copies by a method that assures
numerous identical and durable copies." Most paleontologists have not seen
Pickering's work, since he only sent it out to a few colleages and has not archived
it in libraries. Thus "Walkersaurus" is a nomen nudum. It was
never described in Pickering's 1995 or 1999 publications, though he has stated
it will be described in a future publication- "Mutanda Dinosaurologica".
Much of Pickering's information is derived from Welles' unpublished work on
European theropods, and he/they viewed Duriavenator as a dilophosaurid
along with Liliensternus (DML, 2001, 2002). Holtz (2000) found Duriavenator
to have several possible positions as a tetanurine less derived than Afrovenator
+ Avetheropoda in his cladistic analysis. Most recently, Benson (2008) has redescribed
the material and placed it in a new genus- Duriavenator. Tetanurine characters
he lists include the prominent anterior maxillary process and band-like enamel
grooves. He assigns it to (a stem-based) Spinosauroidea based on a supposed
maxillary fossa, but this seems more likely to be the promaxillary fenestra
based on its position. The more posterior pneumatic recess is then the maxillary
antrum, which opens posteriorly via a fenestra. Yet medially closed promaxillary
fenestrae are probably primitive for avepods, as they are present in Dilophosaurus,
ceratosaurs and Piatnitzkysaurus as well. Benson further referred it
to Megalosauridae+Spinosauridae based on the anteriorly wide paradental groove.
However, Benson et al. (2008) note that Dilophosaurus and Allosaurus
also have open interdental grooves, while Torvosaurus' is completely
closed. Benson (2008) found Duriavenator to emerge as a megalosaurid
in his unpublished phylogenetic analysis.
References- Owen, 1883. On the skull of Megalosaurus. Quarterly
Journal of the Geological Society of London. 39, 334-347.
Walker, 1964. Triassic reptiles from the Elgin area: Ornithosuchus and
the origin of carnosaurs. Philosophical Transactions of the Royal Society of
London B. 248, 53-134.
Waldman, 1974. Megalosaurids from the Bajocian (Middle Jurassic) of Dorset.
Palaeontology. 17, 325-339.
Pickering, 1995. Jurassic Park: Unauthorized Jewish Fractals in Philopatry,
A Fractal Scaling in Dinosaurology Project, 2nd revised printing. Capitola,
California: 478 pp. [January 27, 1995].
Welles and Pickering, 1999. An Extract From: Archosauromorpha: Cladistics and
Osteologies. 70 pp.
Holtz, 2000. A new phylogeny of the carnivorous dinosaurs. Gaia 15. 5-61.
Pickering, DML 2001. http://dml.cmnh.org/2001Dec/msg00510.html
Allain, 2002a. Les Megalosauridae (Dinosauria, Theropoda). Nouvelle découverte
et révision systématique: Implications phylogénétiques
et paléobiogéographiques. Unpublished thesis. 329 pp.
Pickering, DML 2002. http://dml.cmnh.org/2002Jan/msg00608.html
Benson, 2008. A redescription of 'Megalosaurus' hesperis (Dinosauria,
Theropoda) from the Inferior Oolite (Bajocian, Middle Jurassic) of Dorset, United
Kingdom. Zootaxa. 1931, 57-67.
Benson, 2008. A new theropod phylogeny focusing on basal tetanurans and its
implications for European 'megalosaurs' and Middle Jurassic dinosaur endemism.
Journal of Vertebrate Paleontology. 28(3), 51A.
Kaijiangosaurus He, 1984
K. lini He, 1984
Bathonian-Callovian, Middle Jurassic
Xiashaximiao Formation, Sichuan, China
Holotype- (CCG 20020) seven cervical vertebrae
Referred- (CCG coll.) two dorsal vertebrae, seven caudal vertebrae, scapula,
coracoid, humerus, proximal ulna, three metacarpals, three manual unguals, proximal
tibia, proximal fibula, metatarsal II, partial phalanx II-1, phalanx II-2, pedal
ungual II, partial metatarsal III (315 mm), partial phalanx III-2, phalanx III-3,
pedal ungual III, incomplete metatarsal IV, phalanx IV-1, phalanx IV-2, phalanx
IV-3, phalanx IV-4, pedal ungual IV
? (CCG coll.) jugal, teeth, femur (400 mm)
Comments- The material comes from at least two individuals, the femur
being comparatively smaller than the other elements. It is possibly synonymous
with Xuanhanosaurus, "Szechuanoraptor" and/or Gasosaurus.
References- He, 1984. [The Vertebrate Fossils of Sichuan]. Sichuan Scientific
and Technical Publishing House, Chengdu, Sichuan. 168 pp. (In Chinese)
Megalosauri Fitzinger, 1843
Megalosauria Bonaparte, 1850
Megalosauroides Gervais, 1852
Megalosauridae Huxley, 1869
Definition- (Megalosaurus bucklandii <- Spinosaurus aegyptiacus,
Allosaurus fragilis, Passer domesticus) (Holtz et al., 2004)
Other definitions- (Torvosaurus tanneri + Afrovenator abakensis
+ "Poekilopleuron" valesdunensis) (modified from Allain, 2002)
Megalosauroidea Huxley, 1869 sensu Nopcsa, 1928
Megalosaurinae Huxley, 1869 sensu Nopcsa, 1928
Definition- (Megalosaurus bucklandii <- Eustreptospondylus oxoniensis)
(Holtz et al., 2004)
Other definitions- (Poekilopleuron bucklandii <- Torvosaurus tanneri)
(modified from Allain, 2002)
Comments- Traditionally a large paraphyletic family containing most basal
tetanurines and carnosaurs, Megalosauridae has been limited since the late 1980's
to include far fewer taxa, generally those here placed in Torvosauridae and
Eustreptospondylinae, plus a few other genera such as Poekilopleuron.
Similarly, the larger group containing these taxa plus Spinosauridae has often
been referred to as Megalosauroidea (e.g. Holtz, 1995), since Megalosauridae
has priority over both Torvosauridae and Spinosauridae. This phylogenetic placement
has been largely based on torvosaurid-like elements referred to Megalosaurus,
but which may not belong to that genus, as more than one large theropod taxon
is present in the type locality. Based on the lectotype dentary alone, Megalosaurus
could fit most anywhere in basal Tetanurae, or even as a carnosaur or basal
ceratosaur. Unfortunately, as one of the earliest theropods named, Megalosaurus'
eponymous family-level taxa (Megalosauroidea, Megalosauridae, Megalosaurinae)
have priority over any other family level name. So if Megalosaurus ends
up being more closely related to Torvosaurus than to Spinosaurus,
Torvosauridae must be renamed Megalosauridae, and Spinosauroidea must be renamed
Megalosauroidea. Similarily, if Megalosaurus is more closely related
to Eustreptospondylus than to Torvosaurus or Spinosaurus,
Eustreptospondylinae must be renamed Megalosaurinae. These issues will be resolved
as more of the referred Megalosaurus remains are restudied and described.
References- Bonaparte, 1850. Conspectus Systematum Herpetologiae et Amphibiologiae.
Editio Altera Reformata [Survey of the systems of reptiles and amphibians. Second
revised edition]. E. J. Brill, Leyden 1.
Holtz, 1995. A new phylogeny of the Theropoda. Journal of Vertebrate Paleontology.
15(3, suppl.), 35A.
Megalosaurus Buckland, 1824
= "Megalosaurus" Parkinson, 1822
M. bucklandii Matell, 1827
= Megalosaurus "conybeari" Ritgen, 1826
= Metriacanthosaurus "brevis" Welles, Powell and Pickering
vide Pickering 1995
Aalenian-Oxfordian, Middle Jurassic-Late Jurassic
Corallian Oolite Formation?, Taynton Limestone Formation (=Stonesfield Slate),
England
Lectotype- (OUM J13505) anterior dentary, teeth
Paralectotypes- ?(OUM J13560) ilium (703 mm) (Walker, 1964)
?(OUM J13561) femur (740 mm) (Day and Barrett, 2004)
?(OUM J13563) pubis
?(OUM J13565) ischium
?(OUM J13572) metatarsal II
?(OUM J13576) sacrum
?(OUM J13577) posterior dorsal vertebra
?(OUM J13579) proximal caudal vertebra
?(OUM J13580) rib
?(OUM J29792) rib
?(OUM J29881) ilium
?(OUM coll.) several caudal vertebrae
Referred- ?(BMNH 2581) incomplete caudal vertebra (Lydekker, 1888)
?(BMNH 25581) caudal vertebra (Welles and Pickering, 1999)
?(BMNH 25582) proximal pubis (Lydekker, 1888)
?(BMNH 28301) fragmentary ischium (Lydekker, 1888)
?(BMNH 28957) two sacral vertebrae (Lydekker, 1888)
?(BMNH 31808) incomplete femur (815 mm) (Day and Barrett, 2004)
?(BMNH 31809) proximal tibia (Lydekker, 1888)
?(BMNH 31810) coracoid (150 mm) (Lydekker, 1888)
?(BMNH 31811; intended holotype of Metriacanthosaurus "brevis")
partial ilium (Lydekker, 1888)
?(BMNH 31813) two partial dorsal vertebrae (Lydekker, 1888)
?(BMNH 31824) rib (Lydekker, 1888)
?(BMNH 31825) rib (Lydekker, 1888)
?(BMNH 31828) jugal (Welles and Pickering, 1999)
?(BMNH 31834) tooth (Lydekker, 1888)
?(BMNH 31932) pedal phalanx (Lydekker, 1888)
?(BMNH 32725) tibia (740 mm), metatarsus (452 mm)
?(BMNH 36585) sacral rib, ulna (Lydekker, 1888)
?(BMNH 40125a) metatarsal (Lydekker, 1888)
?(BMNH 40131) partial coracoid (Lydekker, 1888)
?(BMNH 41305) tooth (Lydekker, 1888)
?(BMNH 42024) tooth (Lydekker, 1888)
?(BMNH 44097) rib (Lydekker, 1888)
?(BMNH 44097a) rib (Lydekker, 1888)
?(BMNH 47963) tooth (Lydekker, 1888)
?(BMNH 378303) rib (Lydekker, 1888)
?(BMNH R234) tooth (Lydekker, 1888)
?(BMNH R283) partial ilium (Lydekker, 1888)
?(BMNH R285) cervical vertebra (Lydekker, 1888)
?(BMNH R700) sacrum (Lydekker, 1888)
?(BMNH R1098) partial sacrum (Welles and Pickering, 1999)
....(BMNH 29857) partial sactum (Welles and Pickering, 1999)
?(BMNH R1099) (scapcor 830 mm) proximal scapula, partial coracoid (Lydekker,
1888)
?(BMNH R1100) ilium (Lydekker, 1888)
?(BMNH R1101) ilium (Lydekker, 1888)
?(BMNH R1102) tibia (Lydekker, 1888)
?(BMNH R1103) distal tibia (Lydekker, 1888)
?(BMNH R1104) metatarsal (Lydekker, 1888)
?(BMNH coll.) ulna, other elements (Lydekker, 1888)
?(BMNH coll.) tibia (Welles and Pickering, 1999)
?(Royal College of Surgeons coll.) tooth, tooth fragments, femur, partial femur,
tibia, phalanx, bone fragment (Owen, 1954)
?(Duke of Marlborough coll.) partial mandible (lost) (Owen, 1857)
?(GPIT 18392) proximal femur, fibula (Welles and Pickering, 1999)
?(GSM 3887) sacrum (Welles and Pickering, 1999)
?(GSM 109560, paratype of Scelidosaurus harrisoni) femur fragment (Owen
1859)
?(GSM 109561) ungual (Owen 1859)
?(OUM J12142) partial mandible, teeth
?(OUM J13506) maxilla
?(OUM J13562) tibia (Galton and Molnar, 2005)
?(OUM J13567) incomplete pubis
?(OUM J13568) incomplete tibia
?(OUM J13569) metatarsal III
?(OUM J13573) metatarsal II
?(OUM J13574) scapulocoracoid (845 mm) (Walker, 1964)
?(OUM J13575) humerus
?(OUM J13578) proximal caudal vertebra
?(OUM J13598) tooth (Benton and Spencer, 1995)
?(OUM J13599) maxilla
?(OUM J13882) tooth (Benton and Spencer, 1995)
?(OUM J29754) proximal femur
?(OUM J29758) distal tibia
?(OUM J29761) humerus
?(OUM J29765) proximal scapula (Benton and Spencer, 1995)
?(OUM J29773) tooth (Benton and Spencer, 1995)
?(OUM J29797) partial pubis
?(OUM J29803) incomplete femur (735 mm) (Day and Barrett, 2004)
?(OUM J29813) posterior mandible
?(OUM J29872) incomplete ischium
?(OUM J29879) incomplete scapula (765 mm)
?(OUM J29882) ilium
?(OUM J29883) partial ilium
?(OUM J29884) partial ilium
?(OUM J29887a) proximal scapula
?(OUM J29888) (scapcor 795 mm) partial scapula, partial coracoid
?(OUM J29889) partial scapula, partial coracoid
?(OUM coll.) partial dorsal centrum (lost)
?(Phillips coll.) pedal ungual (lost) (Welles and Pickering, 1999)
?(RCS 72) femur (lost) (Welles and Pickering, 1999)
?(RCS 73) partial femur (lost) (Welles and Pickering, 1999)
?(RCS 74) distal tibia (lost) (Welles and Pickering, 1999)
?(SDM 44.23) femur (820 mm) (Day and Barrett, 2004)
?(SMC D.11.34C) partial scapula
? posterior skull fragment (Huene, 1906)
? dorsal vertebra (Huene, 1966)
Diagnosis- (after Benson et al., 2008) longitudinal groove in the ventral
part of the lateral surface of the dentary; slit-like foramen anterior to the
termination of
the Meckelian groove.
Comments- The lectotype dentary was collected in 1797. Parkinson (1822)
used the name "Megalosaurus", but without description or indication
of type material, making it a nomen nudum. The same can be said of Ritgen's
(1826) species Megalosaurus "conybeari". Molnar et al. (1990)
incorrectly attributed the species M. bucklandii to Ritgen, 1826. Meyer
(1832) used the spelling bucklandi for the species, and has been followed
by numerous authors, but this is incorrect.
All paralectotype and referred material is only provisionally referred to the
taxon, since it is either incomparable to the lectotype dentary, or has yet
to be described. There were at least two large theropods present in the Stonesfield
Slate, so at least some of the above material may belong to this second taxon
(= Metriacanthosaurus? "reynoldsi"?; see unnamed abelisaurian
(Day and Barrett, 2004)).
Day and Barrett separated supposed Megalosaurus bucklandii femora into
two groups. Morphotype A is straight in anterior view, slightly anteriorly convex
in lateral view, has a sharp longitudinal ridge anteriorly on the proximal third
of the shaft, and lacks an extensor groove. Morphotype B is strongly sigmoidal
in anterior view, slightly sigmoidal in lateral view, lacks the ridge and has
an extensor groove. One of the paralectotype specimens of M. bucklandii
is morphotype B, so that morphology is provisionally assigned to that taxon.
The desired holotype of Metriacanthosaurus "brevis" is an ilium
illustrated by Day and Barrett, which they couldn't distinguish from M. bucklandii.
Pickering says of M. "brevis" and Metriacanthosaurus parkeri,
"their great height and shortness -- diagnostic for Metriacanthosaurus
-- separates them from ilia of Megalosaurus bucklandii and Allosaurus."
However, Day and Barrett found this was an illusion caused by a broken postacetabular
process and a plaster reconstructed preacetabular process. By using Photoshop
to resize and rotate the M. "brevis" ilium and overlay it on
a M. bucklandii ilium, this is quite apparent. It is not necessarily
shorter, and in fact only differs from M. bucklandii in a few very minor
proportions easily caused by individual variation and/or distortion. Indeed,
the characters Pickering lists as distinguishing M. "brevis"
from M. parkeri are largely also those that distinguish M. bucklandii
from M. parkeri (nearly straight upper margin; taller preacetabular process),
or don't distinguish M. "brevis" from M. bucklandii
(narrower notch between preacetabular process and pubic peduncle; longer, lower
acetabulum; 250 mm long). So I see no reason to support M. "brevis"
and provisionally synonymize it with M. bucklandii. The referred remains
of M. "brevis" include a pectoral girdle Day and Barrett couldn't
distinguish from M. bucklandii, two tibiae I have no information on,
and one of the morphotype A femora. If this femur (or another diagnostic element)
could be shown to be associated with the ilium, my opinion would be revised,
of course. But even in this case, I see no reason to assign the morphotype A
femora to a Metriacanthosaurus, since the latter has a strongly sigmoidal
femur and sinraptorids have deep extensor grooves, medially oriented heads,
and no groove between the lateral condyle and tibiofibular crest.
While Benson et al. (2008) state "There is currently no positive evidence
on which to refer M. bucklandii to Spinosauroidea (including eustreptospondylines
in their opinion) and the characteristic morphology of the dentary that is present
in all known spinosauroids is absent in Megalosaurus", there is
only one character noted in the paper that would exclude Megalosaurus
from the Spinosauroidea- the third alveolus isn't largest (the fourth is). They
further state that in spinosauroids (Torvosaurus, Dubreuillosaurus,
Eustreptospondylus, Magnosaurus, spinosaurids), the dentary is
laterally expanded to accomodate this alveolus, whereas Megalosaurus
shows a much more subtle expansion around the third to sixth alveoli. Yet the
authors also note a character of Megalosaurus which is only found in
some spinosauroids sensu lato- paradental groove open only anteriorly. This
is shared with Dubreuillosaurus and Duriavenator, but not with
Eustreptospondylus, Magnosaurus or Torvosaurus. This leaves
the evidence for referring the lectotype dentary to Spinosauroidea sensu lato
ambiguous.
References- Parkinson, 1822. Outlines of Oryctology. An introduction
to the study of fossil organic remains, especially those found in the British
Strata: London, viii + 350pp.
Buckland, W. 1824. Notice on the Megalosaurus or great fossil lizard
of Stonesfield. Transactions of the Geological Society of London, 21, 390397.
Ritgen, 1826. Versuchte Herstellung einiger Becken urweltlichter Thiere. Nova
Acta Caesareaa Leopold.-Carol. Ger. Nat. Curiosorum. 13, 331-358.
Owen, 1854. Descriptive catalogue of the fossil organic remains of reptilia
and pisces contained in the Museum of The Royal College of Surgeons of England:
184pp.
Owen, 1857. Monograph on the Fossil Reptilia of the Wealden Formations. Part
III. Dinosauria (Megalosaurus). Palaeontographical Society Monographs.
34, 1-26, pls 112.
Phillips, 1871. Geology of Oxford and the Valley of the Thames: Oxford at the
Clarendon Press, 523pp.
Lydekker, 1888. Catalogue of the Fossil Reptilia and Amphibia in the British
Museum (Natural History), Cromwell Road, S.W., Part 1. Containing the Orders
Ornithosauria, Crocodilia, Dinosauria, Squamta, Rhynchocephalia, and Proterosauria:
British Museum of Natural History, London, 309pp.
Huene, 1906. Ueber das Hinterhaupt von Megalosaurus bucklandi aus Stonesfield.
Neues Jahrbuch für Mineralogie, Geologie und Paläontologie, 1906,
p. 1-12.
Huene, 1926. The carnivorous Saurischia in the Jura and Cretaceous formations,
principally in Europe. Revista del Museo de La Plata. 29, 1-167.
Huene, 1966. Ein Megalosauriden-Wirbel des lias aus norddeutschem Geschiebe:
Neues Jahrbuch für Mineralogie, Geologie und Paläontologie, 1966,
n. 5, p. 318-319.
Benton and Spencer, 1995. Fossil Reptiles of Great Britain: Geological Conservation
Review Series, Chapman & Hall, 386pp.
Pickering, 1995. "Jurassic Park: Unauthorized Jewish Fractals in Philopatry,"
A Fractal Scaling in Dinosaurology Project, 2nd revised printing, Capitola,
California: 478 pp. [January 27, 1995].
Welles and Pickering, 1999. Megalosaurus bucklandii: Private publication
of Stephen Pickering, An extract from Archosauromorpha: Cladistics & Osteologies.
A Fractal Scaling in Dinosaurology Project, p. 1-119.
Allain, 2002a. Les Megalosauridae (Dinosauria, Theropoda). Nouvelle découverte
et révision systématique: Implications phylogénétiques
et paléobiogéographiques. Unpublished thesis. 329 pp.
Day and Barrett, 2004. Material Referred to Megalosaurus (Dinosauria:
Theropoda) from the Middle Jurassic of Stonesfield, Oxfordshire, England: one
taxon or two? Proceedings of the Geologists' Association. 115, 359-366.
http://groups.yahoo.com/group/paleo_bio_dinosaur_ontology/message/8460
Galton and Molnar, 2005. Tibiae of small theropod dinosaurs from Southern England,
from the Middle Jurassic of Stonesfield near Oxford and the Lower Cretaceous
of the Isle of Wight: In: The Carnivorous Dinosaurs, Edited by Carpenter, K.,
I. Theropods Old and New, p. 3-22.
Benson, Barrett, Powell and Norman, 2008. The taxonomic status of Megalosaurus
bucklandii (Dinosauria, Theropoda) from the Middle Jurassic of Oxfordshire,
UK. Paleontology. 51, 419-424.
Benson, 2009. An assessment of variability in theropod dinosaur remains from
the Bathonian (Middle Jurassic) of Stonesfield and New Park Quarry, UK and taxonomic
implications for Megalosaurus bucklandii and Iliosuchus incognitus.
Palaeontology. 52(4), 857-877.
Benson, in press. A description of Megalosaurus bucklandii (Dinosauria:
Theropoda) from the Bathonian of the United Kingdom and the relationships of
Middle Jurassic theropods. Zoological Journal of the Linnean Society.
M? phillipsi Welles,
Powell and Pickering vide Pickering, 1995
Kimmeridgian, Late Jurassic
Kimmeridgian Clay, England
Material- (OUM J29886) tibia
....(OUM J13586) metatarsal II, metatarsal III, metatarsal IV
Diagnosis- (from Pickering, DML 2002) compared with Megalosaurus bucklandii,
tibia with more expanded head, the shaft straighter and more massive; the fibular
flange begins higher on the shaft and extends farther down; if this is oriented
laterally, then the head is rotated much more craniolaterally, its axis 55 degrees
versus 40 degrees; the astragalar facet lacks the dorsal pit, but has the same
curvature of the overhang; the concavity of the distal end is wider; the postfibular
plate does not extend so far below the medial end.
Comments- OUM J29886 is intended as the holotype. The validity and referral
of this specimen to Megalosaurus has not been justified in published
literature.
References- Phillips, 1871. Geology of Oxford and the Valley of the Thames.
Clarendon Press, Oxford.
Huene, 1926. The carnivorous Saurischia in the Jura and Cretaceous formations,
principally in Europe. Revista del Museo de La Plata. 29, 35-167.
Pickering, 1995. "Jurassic Park: Unauthorized Jewish Fractals in Philopatry,"
A Fractal Scaling in Dinosaurology Project, 2nd revised printing, Capitola,
California: 478 pp. [January 27, 1995].
http://dml.cmnh.org/2002Mar/msg00553.html
M? sp. indet. (Lydekker, 1888)
Early Jurassic
Coral Rag of Dry Sandford, England
Material- (BMNH R1027) sacrum
Comments- This specimen is probably too early to be Megalosaurus,
and cannot be referred to that taxon because the sacrum of M. bucklandii
lacks described apomorphies and known specimens are only tentatively referred
to the species.
Reference- Lydekker, 1888. Catalogue of the Fossil Reptilia and Amphibia
in the British Museum (Natural History), Cromwell Road, S.W., Part 1. Containing
the Orders Ornithosauria, Crocodilia, Dinosauria, Squamta, Rhynchocephalia,
and Proterosauria: British Museum of Natural History, London, 309pp.
M? sp. indet. (Lydekker, 1888)
Bajocian, Middle Jurassic
Inferior Oolite, England
Material- (BMNH 47152) tooth
(BMNH R497) partial tooth
Comments- These specimens cannot be referred to Megalosaurus bucklandii
because the teeth of that taxon lack described apomorphies.
Reference- Lydekker, 1888. Catalogue of the Fossil Reptilia and Amphibia
in the British Museum (Natural History), Cromwell Road, S.W., Part 1. Containing
the Orders Ornithosauria, Crocodilia, Dinosauria, Squamta, Rhynchocephalia,
and Proterosauria: British Museum of Natural History, London, 309pp.
M? sp. indet. (Lydekker, 1888)
Middle Bathonian, Middle Jurassic
Cotswold Slates, England
Material- (BMNH 28608) tooth (Lydekker, 1888)
(BMNH R2635) tooth (Benton and Spencer, 1995)
(GLCRM T.70-1, G.72-3, T.74-6) ribs, limb elements (Benton and Spencer, 1995)
Comments- These specimens cannot be referred to Megalosaurus bucklandii
because the teeth of that taxon lack described apomorphies and known postcrania
are only tentatively referred to the species.
References- Lydekker, 1888. Catalogue of the Fossil Reptilia and Amphibia
in the British Museum (Natural History), Cromwell Road, S.W., Part 1. Containing
the Orders Ornithosauria, Crocodilia, Dinosauria, Squamta, Rhynchocephalia,
and Proterosauria: British Museum of Natural History, London, 309pp.
Benton and Spencer, 1995. Fossil Reptiles of Great Britain: Geological Conservation
Review Series, Chapman & Hall, 386pp.
M? sp. indet. (Lydekker, 1888)
Late Bathonian, Middle Jurassic
Cornbrash Formation, England
Material- (BMNH 47169) dorsal centrum
Comments- This specimen cannot be referred to Megalosaurus bucklandii
because the vertebrae of that taxon lack described apomorphies and known specimens
are only tentatively referred to the species.
Reference- Lydekker, 1888. Catalogue of the Fossil Reptilia and Amphibia
in the British Museum (Natural History), Cromwell Road, S.W., Part 1. Containing
the Orders Ornithosauria, Crocodilia, Dinosauria, Squamta, Rhynchocephalia,
and Proterosauria: British Museum of Natural History, London, 309pp.
M? sp. indet. (Delair, 1973)
Late Bathonian, Middle Jurassic
Forest Marble, England
Material- (BMNH 39476) tooth (Delair, 1973)
teeth (Freeman, 1979)
Comments- These specimens cannot be referred to Megalosaurus bucklandii
because the teeth of that taxon lack described apomorphies.
References- Delair, 1973. The Dinosaurs of Wiltshire: Whiltshire Archaeology
and Natural History Magazine, v. 68, p. 1-7.
Freeman, 1979. A Middle Jurassic Mammal Bed from Oxfordshire: Palaeontology,
v. 22, part 1, p. 135-166.
M? sp. indet. (Lydekker, 1888)
Early Cretaceous
Potton Sands, England
Material- (BMNH 42028) fragmentary posterior dorsal vertebra
Comments- This specimen is too late to be Megalosaurus, and cannot
be referred to that taxon because the vertebrae of M. bucklandii lack
described apomorphies and known specimens are only tentatively referred to the
species.
Reference- Lydekker, 1888. Catalogue of the Fossil Reptilia and Amphibia
in the British Museum (Natural History), Cromwell Road, S.W., Part 1. Containing
the Orders Ornithosauria, Crocodilia, Dinosauria, Squamta, Rhynchocephalia,
and Proterosauria: British Museum of Natural History, London, 309pp.
M? sp. indet. (Owen, 1854)
Hauterivian-Barremian, Early Cretaceous
Weald Clay, England
Material- (Royal College of Surgeons coll.) posterior dorsal vertebra
(Royal College of Surgeons coll.) dorsal centrum, caudal vertebra, distal caudal
vertebra
(Royal College of Surgeons coll.) partial neural arch
(Royal College of Surgeons coll.) ungual
Comments- These specimens are too late to be Megalosaurus, and
cannot be referred to that taxon because the vertebrae and unguals of M.
bucklandii lack described apomorphies and known specimens are only tentatively
referred to the species.
Reference- Owen, 1854. Descriptive catalogue of the fossil organic remains
of reptilia and pisces contained in the Museum of The Royal College of Surgeons
of England: 184pp.
M? sp. indet. (Sauvage, 1900)
Bathonian, Middle Jurassic
Indre, France
Material- tooth
Comments- This specimen cannot be referred to Megalosaurus bucklandii
because the teeth of that taxon lack described apomorphies.
Reference- Sauvage, 1900. Note sur les poissons et les reptiles du Jurassique
inferieur du department de l'Indre: Bulletin de la societie geologiques de France,
3rd series, v. 28, p. 500-504.
M? sp. indet. (Mercier, 1937)
Early Bathonian, Middle Jurassic
Ecouche Limestone, France
Material- tooth
Comments- This specimen cannot be referred to Megalosaurus bucklandii
because the teeth of that taxon lack described apomorphies.
References- Mercier, 1937.
Buffetaut, Cuny and Le Loeuff, 1991. French Dinosaurs: The best record in Europe?:
In: Special Issue, Dinosaur Studies, Commemorating the 150th Aniversary of Richard
Owens Dinosauria. Modern Geology, v. 16, n. 1 and 2, p. 17-42.
M? sp. indet. (Royo and Gomez, 1927)
Hauterivian-Barremian, Early Cretaceous
Weald Clay, Spain
Comments- These specimens are too late to be Megalosaurus.
Reference- Royo and Gomez, 1927. Sur le facies wealdien d'Espange: Bulletin
de la societie geologiques de France, 4th series, v. 27, CR, p. 125-128.
M? sp. indet. (Ubaghs, 1892)
Late Maastrichtian, Late Cretaceous
Maastricht Formation, Belgium
Material- fragmentary tooth
Comments- This specimen is too late to be Megalosaurus, and cannot
be referred to that taxon because the teeth of M. bucklandii lack described
apomorphies. Based on its age, this is more probably abelisaurid or dromaeosaurid.
Reference- Ubaghs, C., 1892, Le Megalosaurus dans la craie superieure
du Limbourg: Bull. Soc. Gelge. Geol., v. 6, Mem, p. 26-29.
Merosaurus Welles, Powell
and Pickering vide Pickering, 1995
M. newmani Welles, Powell and Pickering vide Pickering, 1995
Sinemurian, Early Jurassic
Lower Lias, England
Material- (BMNH 39496, holotype of Scelidosaurus harrisoni) distal
femur, proximal tibia
(GSM 109560; lost) femur, partial tibia
?(GSM 109561) manual ungual I
Description- (after Pickering, online 2005) the type is very similar
to Sarcosaurus, and is of the same diagnostic pattern as Ceratosaurus
and Dilophosaurus in the nearly straight cranial edge, the concave popliteal
notch, and the elliptical, caudolaterally oriented ectocondyle. The anterior
trochanter is a blunt cone vs. the diagnostic flat blade of Megalosauridae.
Comments- This was originally referred to Scelidosaurus by Owen
(1863; 1884), and was designated as the type of that taxon by Lydekker (1888).
Newman (1968) recognized its theropod nature. Carrano and Sampson (2004) refer
this specimen to the basal Tetanurae.
References- Pickering, S., 1995. "Jurassic Park: Unauthorized Jewish
Fractals in Philopatry," A Fractal Scaling in Dinosaurology Project, 2nd
revised printing, Capitola, California: 478 pp. [January 27, 1995].
Carrano and Sampson, 2004. A review of coelophysoids (Dinosauria: Theropoda)
from the Early Jurassic of Europe, with comments on the late history of the
Coelophysoidea. N. Jb. Geol. Palaont. Mh. 2004 (9): 537-558.
http://groups.yahoo.com/group/paleo_bio_dinosaur_ontology/message/8446
"Mifunesaurus" Hisa, 1985
Middle Cenomanian, Late Cretaceous
Kabu Formation of the Mifune Group, Japan
Material- (YNUGI 10003; Mifune-ryu) (~6-7 m) maxillary tooth (72.7 mm)
Comments- This tooth was discovered in 1979, and described in 1984 by
Hawegawa and Murata. They referred it to Megalosauridae gen. et. sp. indet.,
and nicknamed it Mifune-ryu. Hisa (1985) later gave it the nomen nudum
"Mifunesaurus" in an illustrated booklet on dinosaurs. It was illustrated
and described in detail by Hasegawa et al. (1992), who referred it to Megalosauridae
based on similarity to "Walkersaurus", Gasosaurus and a Chinese
specimen of unknown stratigraphic origin probably incorrectly referred to Megalosaurus
bucklandii. They thought it was distinct from other named theropods in its
low basal width to crown length ratio (.17), but this is also seen in ceratosaurids,
for instance. It remains a nomen nudum because Hawegawa et al. did not
name the tooth, while Hisa's (1985) description was apparently deficient. While
Chure (2000) stated some of the teeth from the Kabu Formation have enamel wrinkles,
Chure et al. (1999) show he was referring to a tooth from the Jobu Formation
(MDM 341). Additional remains from the Jobu Formation of the Mifune Group (Tamara
et al., 1991- four teeth, tibia, fibula, metatarsals II and III) are sometimes
referred to "Mifunesaurus" as well, but although Chure (2000) stated
the teeth were similar, there are no reported synapomorphies which would allow
placing them in the same taxon.
The tooth crown is 72.7 mm tall, with a FABL of 22.5 mm and a basal width of
12.3 mm. It is recurved and lens shaped in section, with almost symmetrically
distributed carinae. There are 20 serrations per 5 mm on both mesial and distal
carinae. Blood grooves are present at least posteriorly, but enamel ridges are
absent. "Mifunesaurus"' tooth seems too thick to be a ceratosaurid,
and taller than in abelisaurids, so is probably from a non-maniraptoriform tetanurine,
such as a spinosauroid or carnosaur.
References- Hasegawa and Murata, 1984. First Record of Carnivorous Dinosaur
from the Upper Cretaceous of Kyushu, Japan. Abstract of the Annual Meeting of
the Paleontological Society of Japan.
Hisa, 1985.
Dong, Hasegawa and Azuma, 1990. The Age of Dinosaurs in Japan and China. Fukui,
Japan: Fukui Prefectural Museum. 65 pp.
Tamara, Okazaki and Ikegami, 1991. Occurence of carnosaurian and herbivorous
dinosaurs from upper formation of Mifune Group, Japan, Memiors of the Faculty
of Education, Kumamoto University. 40, 31-45.
Hasegawa, Murata, Wasada and Manabe, 1992. The first carnosaur (Saurischia;
Theropoda) from Japan; A tooth from the Cenomanian Mifune Group of Kyushu. Sci.
Rep. Yokohama Natl. Univ. Ser. 2 39, 41-49.
Chure, Manabe, Tanimoto and Tomida, 1999. An unusual theropod tooth from the
Mifune Group (Late Cenomanian to Early Turonian), Kumamoto, Japan. in Tomida,
Rich, and Vickers-Rich (eds.). Proceedings of the Second Gondwanan Dinosaur
Symposium. National Science Museum (Tokyo) Monographs. 15, 291-296.
Chure, 2000. A new species of Allosaurus from the Morrison Formation
of Dinosaur National Monument (Utah-Colorado) and a revision of the theropod
family Allosauridae. Ph.D. dissertation, Columbia University, 1-964.
"Saltriosaurus" Dalla Vecchia,
2001
Sinemurian, Early Jurassic
Saltrio Formation, Italy
Material- (MSNM V3664) (8 m; 1.5 tons) lateral tooth, dorsal rib fragments,
scapular fragment, furcula, humeri, metacarpal II, phalanx II-1, phalanx III-1,
phalanx III-2, manual ungual III, proximal fibula, distal tarsal III, distal
tarsal IV
References- Dalla Vecchia, 2001. A new theropod dinosaur from the Lower
Jurassic of Italy, Saltriosaurus. Dino Press. 3, 8187.
Del Sasso, 2003. Dinosaurs of Italy. Comptes Rendus Palevol.
Shidaisaurus Wu, Currie, Dong,
Pan and Wang, 2009
S. jinae Wu, Currie, Dong, Pan and Wang, 2009
Aalenian-Bajocian, Middle Jurassic
Upper Lufeng Formation, Yunnan, China
Holotype- (LDM-LCA 9701-IV) braincase, three teeth, atlantal intercentrum,
axis, cervical vertebrae, third dorsal vertebra, fourth dorsal vertebra, fifth
dorsal vertebra, sixth dorsal vertebra, seventh dorsal vertebra, eighth dorsal
vertebra, ninth dorsal vertebra, tenth dorsal vertebra, eleventh dorsal vertebra,
twelfth dorsal vertebra, thirteenth dorsal vertebra, two gastralia, sacral vertebrae
1+2, third sacral vertebra, fourth sacral vertebra, fifth sacral vertebra, first
caudal vertebra, second caudal vertebra, third caudal vertebra, ilium (620 mm),
pubes (620 mm), ischium (599 mm)
Diagnosis- (after Wu et al., 2009) supraoccipital excluded from foramen
magnum; paroccipital processes downturned at 110 degrees; large, pointed axial
epipophyses; thin and broad lamina between axial neural spine and epipophyses;
iliopubic ratio ~1.00; obturator notch absent on ischium; ischium >96% of
pubic length.
Comments- Wu et al. merely considered Shidaisaurus a tetanurine,
perhaps non-avetheropod due to the absent axial pleurocoel. Inclusion in my
theropod supermatrix suggests it is either related to basal tetanurines such
as Piatnitzkysaurus and "Szechuanoraptor", or is a basal carnosaur
Reference- Wu, Currie, Dong, Pan and Wang, 2009. A new theropod dinosaur
from the Middle Jurassic of Lufeng, Yunnan, China. Acta Geologica Sinica. 83(1),
9-24.
Valdoraptor Olshevsky, 1991
V. oweni (Lydekker, 1889) Olshevsky, 1991
= Megalosaurus oweni Lydekker, 1889
= Altispinax oweni (Lydekker, 1889) Huene, 1923
Barremian, Early Cretaceous
Upper Weald Clay, England
Holotype- (BMNH R2559) incomplete metatarsal II, incomplete metatarsal
III, incomplete metatarsal IV
Comments- This was originally referred to Hylaeosaurus by Owen,
but was identified as theropod by Hulke (1881).
Lydekker (1890) referred BMNH R604a, 604c-d and 1525 from the earlier Wadhurst
Clay and BMNH 2574, 2661 and 2680 from the type horizon to the species, but
they have not been compared in detail (see Altispinax? sp. indet. entry).
References- Hulke, 1881.
Lydekker, 1889. On the remains and affinities of five genera of Mesozoic reptiles.
Q. J. Geol. Soc. London 45: 41-59.
Lydekker, 1890.
Huene, 1923. Carnivorous Saurischia in Europe since the Triassic. Bull. Geol.
Soc. Am. 34: 449-458.
Olshevsky, 1991."A Revision of the Parainfraclass Archosauria Cope, 1869,
Excluding the Advanced Crocodylia," Mesozoic Meanderings #2 (1st printing):
iv + 196 pp.
unnamed tetanurine (Ricqles, 1967)
Aptian-Albian, Early Cretaceous
Elrhaz Formation of the Tegama Group, Niger
Material- partial manual ungual I
Comments- This specimen was described by Ricqles (1967) as dinosaurian,
and Rozhderstveksky (1970) placed it in Theropoda. Nessov (1995) referred it
to Therizinosauria or "groups most closely related to them" (which
in his opinion consisted of spinosaurids and dryptosaurids). The ungual closely
resembles both Baryonyx and Alxasaurus, though it tapers more
than the former and is less curved than the latter.
References- Ricqles, 1967. La paleontologie de terrain: Un bilan international.
Atomes. 243, 337-341.
Rozhdestvensky, 1970. Giant claws of enigmatic Mesozoic reptiles. Paleontological
Journal. 1970(1), 131-141.
Nessov, 1995. Dinosaurs of nothern Eurasia: new data about assemblages, ecology,
and paleobiogeography. Institute for Scientific Research on the Earth's Crust,
St. Petersburg State University, St. Petersburg. 1-156.
unnamed tetanurine (He, 1984)
Bathonian-Callovian, Middle Jurassic
Xiashaximiao Formation
Material- three cervical vertebrae, two anterior dorsal vertebrae, proximal
caudal vertebra, coracoid, ischium, tibia, fibula, metatarsal
Comments- This specimen was originally referred to Szechuanosaurus,
but was found to be an indeterminate basal tetanurine by Chure (2000).
References- He, 1984. The Vertebrate Fossils of Sichuan. Sichuan Scientific
and Technical Publishing House, Chengdu, Sichuan. 168 pp.
Li, Zhang and Cai, 1998. The Characteristics of the composition of the trace
elements in Jurassic Dinosaur Bones and Red Beds in Sichuan Basin: Geological
Publishing House, Beijing, 155pp.
Chure, 2000. A new species of Allosaurus from the Morrison Formation
of Dinosaur National Monument (Utah-Colorado) and a revision of the theropod
family Allosauridae. Ph.D. dissertation, Columbia University, 1-964.
unnamed tetanurine (Galton and Molnar, 2005)
Middle Jurassic
Tauton Limestone Formation (Stonefield Slate), England
Material- distal tibia
Comments- Assigned to the basal Tetanurae.
Reference- Galton and Molnar, 2005. Tibiae of small theropod dinosaurs
from Southern England: From the Middle Jurassic of Stonesfield near Oxford and
the Lower Cretaceous of the Isle of Wight. In Carpenter (ed.). The Carnivorous
Dinosaurs. Indiana University Press. 3-22.
undescribed tetanurine (Naish and Martill, 2007)
Kimmeridgian, Late Jurassic
Kimmeridge Clay, England
Material- cervical vertebrae, dorsal vertebrae, sacral vertebrae, caudal
vertebrae, pelvic elements, hindlimb elements
Comments- This specimen was mentioned as a "peculiar, gracile tetanurine"
by Naish (online, 2006) and is due to be studied. Naish and Martill (2007) mention
it in print as pers. comm. from Powell.
References- Naish, online 2006. http://darrennaish.blogspot.com/2006/12/obscure-dinosaurs-of-kimmeridge-clay.html
Naish and Martill, 2007. Dinosaurs of Great Britain and the role of the Geological
Society of London in their discovery: basal Dinosauria and Saurischia. Journal
of the Geological Society, London. 164, 493-510.
"Allosaurus" tendagurensis
Janensch, 1925
= Antrodemus tendagurensis (Janensch, 1925) Huene, 1932
Kimmeridgian, Late Jurassic
Middle Saurian Beds of Tendaguru Formation, Tanzania
Holotype- (HM 67) partial tibia (~910 mm)
Diagnosis- combination of- cnemial crest not strongly curved laterally
(basal); no proximal groove on posterior tibial surface separating lateral and
medial condyles (basal); fibular crest distally placed (derived).
Comments- Chure (2000) notes this specimen resembles abelisaurids because
it lacks a strongly curved cnemial crest or incisura tibialis and has no posterior
groove between the lateral and medial condyles. However, he states the astragalus
is not fused to it and the fibular crest is more distally placed than abelisaurids.
The lack of a well developed incisura tibialis indicates
it is not tetanurine, but the distally placed fibular crest is a tetanurine
synapomorphy. The lack of fusion is inconsequential, as ceratosaurs like Elaphrosaurus
and Quilmesaurus exhibit the condition.
References- Janensch, 1925. Die Coelurosaurier und Theropoden der Tendaguru-Schichten
Deutsch-Ostafrikas. Palaeontographica (Suppl. 7)1:1-99.
Huene, 1932. Die fossile Reptile-Ordnung Saurischia, ihre Entwicklung und Geschichte.
Monogr. Geol. Palaeontol. (Pt. I and II, Ser. I) 4, 1-361.
Chure, 2000. A new species of Allosaurus from the Morrison Formation
of Dinosaur National Monument (UT-CO) and a revision of the theropod family
Allosauridae. PhD dissertation. Columbia University. 964 pp.
unnamed tetanurine (Stromer, 1934)
Cenomanian, Late Cretaceous
Baharija Formation, Egypt
Material- (IPHG 1912 VIII 192) tibia (660+ mm) (Stromer 1934)
(IPHG 1912 VIII 76) tibia (575 mm) (Stromer 1934)
(IPHG 1912 VIII 190) tibia (Stromer 1934)
Cenomanian, Late Cretaceous
Kem Kem Beds, Morocco
tibia (630 mm) (Lavocat 1954)
Comments- Stromer described three tibiae from the Baharija Formation
of Egypt and assigned them to Elaphrosaurus. The distally placed fibular
crest and high ascending process suggest this is a tetanurine, but the lack
of a prominent incisura tibialis is more primitive than other tetanurines except
"Allosaurus" tendagurensis and Condorraptor. Lavocat
(1954) described a tibia from the Kem Kem Beds of Morocco that he thought was
very similar to Stromers material, but had a differently developed cnemial
crest.
References- Stromer, 1934. Wirbeltierreste der Baharije-Stufe (unterstes
Cenoman). 13. Dinosauria. Abh. Bayer. Akad. Wissensch. Math.-naturwiss. Abt.
22:1-79.
Lavocat, 1954. Sur les Dinosauriens du continental intercalaire des Kem-Kem
de la Daoura. C. R. 19th Internatl. Geol. Congr. 1952: 65-68.
unnamed basal tetanurine (Rauhut, 2005)
Late Kimmeridgian, Late Jurassic
Middle Saurian Bed of Tengaguru Formation, Tanzania
Material- (MB.R 1763) tibia (163 mm)
Comments- Originally described as 'coelurosaurier A', Rauhut (2005) has
identified it as a basal tetanurine.
References- Janensch, 1925. Die Coelurosaurier und Theropoden der Tendaguru-Schichten
Deutsch-Ostafrikas. Palaeontographica, Supplement VII, 199.
Rauhut, 2005. Post-cranial remains of coelurosaurs (Dinosauria,
Theropoda) from the Late Jurassic of Tanzania. Geol. Mag. 142 (1), pp. 97107.
Condorraptor Rauhut, 2005
C. currumili Rauhut, 2005
Callovian, Middle Jurassic
Canadon Asfalto Formation, Argentina
Holotype- (MPEF-PV 1672) tibia
Paratypes- .........(MPEF-PV 1673) ?fourth cervical vertebra (57 mm)
.........(MPEF-PV 1674) ?seventh cervical vertebra (70 mm)
.........(MPEF-PV 1675) tenth cervical vertebra (59 mm)
.........(MPEF-PV 1676) first dorsal centrum (61 mm)
.........(MPEF-PV 1677) mid dorsal centrum (~70 mm)
.........(MPEF-PV 1678) mid dorsal centrum (~68 mm)
.........(MPEF-PV 1679) partial anterior dorsal neural arch
.........(MPEF-PV 1680) posterior dorsal vertebra (80 mm)
.........(MPEF-PV 1681) second sacral vertebra (75 mm), third sacral vertebra
(75 mm), fourth sacral vertebra (72 mm)
.........(MPEF-PV 1682) mid caudal vertebra (77 mm)
.........(MPEF-PV 1683) distal caudal vertebra (67 mm)
.........(MPEF-PV 1684) dorsal rib fragment
.........(MPEF-PV 1685) dorsal rib fragment
.........(MPEF-PV 1686) ilial fragment
.........(MPEF-PV 1687) ilial fragment
.........(MPEF-PV 1688) pubic fragment
.........(MPEF-PV 1689) partial ischium
.........(MPEF-PV 1690) distal femur
.........(MPEF-PV 1691) distal femur
.........(MPEF-PV 1692) metatarsal IV (242 mm)
.........(MPEF-PV 1693) proximal pedal ungual
.........(MPEF-PV 1694) lateral tooth (FABL 11 mm)
.........(MPEF-PV 1695) lateral tooth (FABL 10.5; ~26 mm)
.........(MPEF-PV 1696) proximal pubis
.........(MPEF-PV 1697) ?second dorsal vertebra (57.5 mm)
.........(MPEF-PV 1700) posterior dorsal vertebra (76 mm)
.........(MPEF-PV 1701) first sacral vertebra (69 mm)
.........(MPEF-PV 1702) anterior caudal vertebra (56 mm)
.........(MPEF-PV 1703) partial chevron
.........(MPEF-PV 1704) ilial fragment
.........(MPEF-PV 1705) anterior dorsal vertebra (65 mm)
Diagnosis- (from Rauhut, 2005) posterior incision between fibular condyle
and medial part of proximal tibia absent; large, shallow depression laterally
on the base of the cnemial crest; pleurocoel in anterior cervical vertebrae
placed behind the posteroventral corner of the parapophyses; large nutrient
foramina on the lateral side of the ischial peduncle of the ilium; metatarsal
IV with a distinct step dorsally between shaft and distal articular facet.
References- Rauhut, 2002. Dinosaur evolution in the Jurassic: a South
American perspective. Journal of Vertebrate Paleontology. 22, 89A.
Rauhut, 2005. Osteology and relationships of a new theropod dinosaur from the
Middle Jurassic of Patagonia. Palaeontology. 48(1), 87-110.
Erectopodidae Huene, 1932
Erectopus Huene, 1923
E. superbus (Sauvage, 1882b) Huene, 1923
= Megalosaurus superbus Sauvage, 1882b
= gen. indet. superbus (Sauvage, 1882b) Huene, 1932
= Erectopus sauvagei Huene, 1932
Early Albian, Early Cretaceous
Phosphate bearing beds of La Penthieve, Meuse, France
Lectotype- (MNHN 2001-4) anterior maxilla
Plastotype- (MNHN 2001-4; holotype of Erectopus sauvagei) incomplete
phalanx I-1 (35 mm), partial manual ungual I, incomplete phalanx II-I, phalanx
II-2, partial manual ungual II, distal metacarpal III, phalanx III-1, proximal
phalanx III-2, femur (480 mm), proximal tibia, distal tibia, calcaneum, metatarsal
II (230 mm)
Syntypes- (lost) two teeth, several dorsal centra (55, 65 mm), dorsal
ribs, partial sacrum, several caudal vertebrae (75 mm), distal radius, distal
ulna, metacarpal I, phalanx III-3, metacarpal IV, partial ilium, proximal fibula
Comments- Barrois (1875) and Sauvage (1876, 1882a) first described two
teeth as Megalosaurus, and Sauvage (1882b) later described them, a jaw
fragment and postcranial material as Megalosaurus superbus. Huene (1923)
separated it from Megalosaurus as the new genus Erectopus superbus.
Sauvage (1882b) did not designate any material as a holotype, and it is not
certain it all derives from one individual. In particular, a much larger distal
femur was found in the same area, and the teeth were seemingly discovered before
the postcrania and maxillary fragment. Huene (1932) incorrectly believed Sauvage
based the name superbus on the original teeth, and as he thought these
were too large to go with the postcrania, he separated the latter as Erectopus
sauvagei. However, there is no evidence Sauvage intended to base superbus
on the teeth, and the fact he waited until describing the postcrania in 1882
to name the species suggests the opposite. Furthermore, Allain (2005) notes
the original tooth matches the preserved maxillary teeth in size. Huene retained
the teeth as gen. indet. superbus, which he believed was an allosaurid
in contrast to Erectopus, which he placed in a monotypic new family.
Allain made the maxillary fragment the lectotype of Erectopus superbus,
as the rest of the material is lost (though some elements are preserved as casts
to form a plastotype). As the tooth matches that on the lectotype, and the rest
of the specimen is no more certainly associated than the maxilla and tooth are
with it, there is no reason to separate the tooth from the postcrania taxonomically.
Notably, the ICZN does not allow a species to lack a genus, and the genus Erectopus
must be attached to the species superbus based on Huene (1923), regardless
which elements each name is based on. Erectopus sauvagei is thus an objective
junior synonym of Erectopus superbus.
Huene (1926) reidentified several elements from Sauvages (1882) description.
The clavicle, and metacarpals are a scapula and distal radius and ulna respectively.
However, it seems the scapular identification is in error, and the element is
actually the posteroventral portion of an ilium (Chure, 2000). Metacarpal I
was originally identified as a manual phalanx (Sauvage, 1882b) and pedal phalanx
IV-? (Huene, 1926), until correctly identified by Molnar (1990). Molnar (1990)
questioned the identification of a metacarpal V by Huene (1926). It is here
identified as a metacarpal IV. A manual phalanx here identified as III-3 was
called a pedal phalanx I-1 by Huene (1926) and a manual phalanx III-1 by Chure
(2000). Allain (2002a) reidentified the posterior mandible as an anterior maxilla.
Several other remains have been referred to Erectopus or Megalosaurus
superbus in the past. Simionescu (1913) compared a tooth from earlier sediments
in Romania to Megalosaurus superbus. Huene (1926) doubted it was the
same species due to age differences, but did continue to call it Erectopus
aff. superbus. It does not seem diagnostic past Neotheropoda.
Huene (1926) referred material from Grandpre (two teeth, proximal metacarpal(?),
two metatarsals), Ber-le-Duc (pedal phalanx II-1) and Blacourt (distal femur)
to Erectopus superbus. He later referred at least the Grandpre postcrania
to his new species Erectopus sauvagei, but retained the teeth in superbus
(which he did not refer to a genus at the time). The teeth and metatarsals were
originally described by Barrois (1875) and Sauvage (1876, 1882a, 1882b) as Megalosaurus.
The proximal metacarpal was illustrated by Sauvage (1882b) and noted as a distal
fibula by Huene, but later reidentified by Chure (2000). The distal femur was
originally described by Sauvage (1876), but determined by Chure to be crocodilian.
None of the theropod remains seem particularly diagnostic and are here excluded
from Erectopus.
Lapparent and Zbyszewski (1957) referred two tooth fragments from the Aptian
of Portugal to Megalosaurus superbus, but these are undiagnostic.
Allain (2002a, b) found Erectopus to emerge as a carnosaur in an unpublished
phylogenetic analysis based on his thesis, which was also the conclusion of
his 2005 redescription of the material. This was based solely on the interpretation
of the distal tibia as indicating a well developed posterior astragalar ascending
process. However, a similar morphology is also seen in the tibia of Poekilopleuron,
which lacks a posterior ascending process. My own inclusion of Erectopus
in a theropod supermatrix suggests it is a more basal tetanurine instead, based
on such characters as the weakly curved manual ungual I, limited contact between
metacarpals I and II, possible presence of manual phalanx IV-1, ventrally directed
femoral head.
References- Barraois, 1975. Les reptiles du terrain Crétacé
du nord-est du Bassin de Paris. Bulletin scientifique, historique et littéraire
du Nord. 6, 1-11.
Sauvage, 1876. Notes sur les reptiles fossiles. Bulletin de la Société
Géologique de France. 4, 435-442.
Sauvage, 1882a. Sur les reptiles trouves dans le Gault de l'est de la France.
Comptes Rendus de l'Académie des Sciences. 94, 1265.
Sauvage, 1882b. Recherches sur les reptiles trouves dans le Gault de l'est du
bassin de Parts. Mémoires de la Société Géologique
de France. 2, 1-42.
Simionescu, 1913. Megalosaurus aus der Unterkreide der Dobrogea. Chi.
Min. Geol. Palaeontol. 1913, 686-687.
Huene, 1923. Carnivorous Saurischia in Europe since the Triassic. Bulletin of
the Geological Society of America. 34, 449-458.
Huene, 1926. The carnivorous Saurischia in the Jura and Cretaceous formations,
principally in Europe. Revista Museo de La Plata. 29, 35-167.
Huene, 1932. Die fossile Reptil-Ordnung Saurischia, ihre Entwicklung und Geschichte.
Monographien zur Geologie und Palaeontologie. 4(1), 361 pp.
Lapparent and Zbyszewski, 1957. [The dinosaurs of Portugal]. Services Geologiques
du Portugal. Memoire 2, 1-63.
Molnar, 1990. Problematic Theropoda: "Carnosaurs". in Weishampel,
Dodson and Osmolska (eds.). The Dinosauria. University of California Press,
Berkeley, Los Angeles, Oxford. 306-317.
Chure, 2000. A new species of Allosaurus from the Morrison Formation
of Dinosaur National Monument (Utah-Colorado) and a revision of the theropod
family Allosauridae. Ph.D. dissertation, Columbia University, 1-964.
Allain, 2002a. Les Megalosauridae (Dinosauria, Theropoda). Nouvelle découverte
et révision systématique: Implications phylogénétiques
et paléobiogéographiques. Unpublished thesis. 329 pp.
Allain, 2002b. The phylogenetic relationships of Megalosauridae within basal
tetanurine theropods (Dinosauria). Journal of Vertebrate Paleontology. 22(3),
31A.
Allain, 2005. The enigmatic theropod dinosaur Erectopus superbus (Sauvage,
1882) from the Lower Albian of Louppy-le-Ch'teau (Meuse, France). In Carpenter
(ed.). The Carnivorous Dinosaurs. Indiana University Press. 72-86.
E? sp. indet. (Stromer, 1934)
Early Cenomanian, Late Cretaceous
Baharija Formation, Egypt
Material- (IPHG 1912 VIII 78) tibia
(IPHG 1912 VIII 85) femur (800 mm)
(IPHG 1912 VIII 190) (juvenile) femur
Reference- Stromer, 1934. Ergebnisse der Forschungsreisen Prof. E. Stromers
in den Wüsten Ägyptens. II. Wirbeltierreste der Baharîje-Stufe
(unterstes Cenoman). 13. Dinosauria. Abh. Bayer. Akad. Wiss., Math.-Nat. Abt.,
(n. s.) 22, 1-79.
Piatnitzkysaurus Bonaparte,
1979
P. floresi Bonaparte, 1979
Callovian, Middle Jurassic
Canadon Asfalto Formation, Argentina
Holotype- (PVL 4073) (4.24 m, 275 kg (Mazzetta et al., 2000 estimated
450 kg)) maxilla, frontal, braincase, anterior dentaries, axis, cervical vertebrae,
dorsal vertebrae, two proximal caudal vertebrae, scapulae, coracoids, humerus,
ulna, partial ilium, incomplete pubis, incomplete ischia, femora (552 mm), tibiae
(492 mm), fibulae
Paratype- (MACN CH 895) incomplete maxilla, two posterior dorsal vertebrae,
two dorsal centra, four sacral vertebrae, humerus, proximal pubis, ischium,
tibia, metatarsal II (253 mm), metatarsal III (290 mm), metatarsal IV (252 mm)
Diagnosis- (after Rauhut, 2004) parasphenoid recess; parasphenoid recesses
communicate; basipterygoid recesses longer anteroposteriorly than high dorsoventrally;
width of the articular surface of the basipterygoid processes is more than twice
their length and the transverse span between the processes of the left and right
side is less than the width of the basal tubera (also in Piveteausaurus?).
References- Bonaparte, 1979. Dinosaurs: A Jurassic assembalge from Patagonia.
Science 205: 1377-1379
Bonaparte, 1986. Les Dinosaures (Carnosaures, Allosauridés, Sauropodes,
Cétiosauridés) du Jurassique moyen de Cerro Cóndor (Chubut,
Argentine). [The Middle Jurassic dinosaurs (carnosaurs, allosaurids, sauropods,
cetiosaurids) from Cerro Cóndor (Chubut, Argentina).] [in French, with
Spanish summ.]. Annales de Paléontologie. Paris, France 72 p. 247-289.
Rauhut, 2004. Braincase structure of the Middle Jurassic theropod dinosaur Piatnitzkysaurus.
Can. J. Earth Sci. 41: 11091122.
Streptospondylidae Kurzanov, 1989
Streptospondylus Meyer, 1832
= "Streptospondylus" Meyer, 1830
S. altdorfensis Meyer, 1832
= Streptospondylus rostromajor Owen, 1842
= Laelaps gallicus Cope, 1867
= Poekilopleuron gallicum (Cope, 1867) Cope, 1869
= Dryptosaurus gallicus (Cope, 1867) Olshevsky, 1991
Late Callovian-Early Oxfordian, Middle Jurassic-Late Jurassic
Vaches Noires Cliffs, Calvados, France
Lectotypes- (MNHN 8605) (subadult) distal pubis
(MNHN 8606) distal fibula
(MNHN 8607) distal tibia (140 mm wide)
(MNHN 8608) astragalus (114 mm wide)
(MNHN 8609) calcaneum
(MNHN 8787) tenth cervical vertebra, first dorsal vertebra (76 mm), second dorsal
vertebra (64 mm), proximal dorsal rib
(MNHN 8788) posterior part of fifth sacral vertebra, first caudal vertebra
(MNHN 8789) fifth or sixth dorsal vertebra (74 mm)
(MNHN 8789) posterior dorsal vertebra (97 mm)
(MNHN 8793) fourth or fifth dorsal vertebra (72 mm)
(MNHN 8794) postzygopophysis of twelfth dorsal vertebra, posterior half of thirteenth
dorsal vertebra, first sacral vertebra (98 mm), anterior part of second sacral
vertebra
(MNHN 8907) three posterior dorsal vertebrae (90, 95 mm)
Referred- (MNHN 9645) distal femur
Diagnosis- (from Allain, 2001) two hypapophyses on anterior dorsal vertebrae;
anterior dorsal centra strongly opisthocoelous and strongly flattened; posterior
dorsal vertebrae platycoelous; elongate mid and posterior dorsal centra; lateral
extension of the medial buttress above the dorsomedial edge of the ascending
process of the astragalus doesnt reach the median part of the distal end
of the tibia; large depression at the base of the ascending process of the astragalus;
lack of posteromedial process on the astragalus.
Comments- This taxon was recently suggested to be the sister taxon of
Eustreptospondylus (Allain, 2001; Smith et al., 2007) based on the presence
of carotid processes in their anterior dorsal vertebrae. Yet analysis of a larger
dataset including all of Smith et al.'s data suggests it may be a basal tetanurine
instead, based on the shallow groove across the astragalar condyles.
References- Meyer, 1830.
Meyer, 1832. Paleologica zur Geschichte der Erde. Frankfurt am Main, 560 p.
Owen, 1842. Report on British fossil reptiles. Report of the British Association
for the Advancement of Science 11: 60-204.
Cope., 1867. Account of extinct reptiles which approach birds. Proceedings of
the Academy of Natural Sciences of Philadelphia: 234-235.
Cope, 1869.
Olshevsky, 1991."A Revision of the Parainfraclass Archosauria Cope, 1869,
Excluding the Advanced Crocodylia," Mesozoic Meanderings #2 (1st printing):
iv + 196 pp.
Allain, 2001. Redescription of Streptospondylus altdorfensis, Cuviers
theropod dinosaur, from the Jurassic of Normandy, Geodiversitas. 23(3), 349-367.
Allain, 2002a. Les Megalosauridae (Dinosauria, Theropoda). Nouvelle découverte
et révision systématique: Implications phylogénétiques
et paléobiogéographiques. Unpublished thesis. 329 pp.
Smith, Makovicky, Hammer and Currie, 2007. Osteology of Cryolophosaurus ellioti
(Dinosauria: Theropoda) from the Early Jurassic of Antarctica and implications
for early theropod evolution. Zoological Journal of the Linnean Society. 151,
377-421.
"Szechuanoraptor"
Chure, 2000
"S." "yandonensis" (Dong, Chang, Li and Zhou, 1978)
new comb.
= Szechuanosaurus "yandonensis" Dong, Chang, Li and Zhou, 1978
= Metriacanthosaurus "carpenteri" Paul, 1988
= Szechuanoraptor dongi Chure, 2000
Oxfordian, Late Jurassic
Shangshaximiao Formation, Sichuan, China
Material- (CV 00214; intended holotype of "Szechuanoraptor dongi")
(3.8 m, 130 kg) axis (55 mm), third cervical vertebra (60 mm), fourth cervical
vertebra (65 mm), fifth cervical vertebra (55 mm), sixth cervical vertebra (65
mm), seventh cervical vertebra (70 mm), eighth cervical vertebra (62 mm), first
dorsal vertebra (60 mm), second dorsal vertebra (60 mm), third dorsal vertebra
(74 mm), fourth dorsal vertebra (60 mm), seventh dorsal vertebra (65 mm), eighth
dorsal vertebra (62 mm), ninth dorsal vertebra (62 mm), tenth dorsal vertebra,
dorsal ribs, gastralia, sacral centrum, eighteen caudal vertebrae, scapula (500
mm), coracoid, humeri (270, 265 mm), ulnae, radii, two distal carpals, metacarpal
I (50 mm), metacarpal II (90 mm), metacarpal III, manual phalanges, manual ungual
I, manual II, manual ungual III, ilium, pubes (~460 mm), ischium (420 mm), femur
(585 mm), tibiae (580 mm), fibula (560 mm), astragali, calcanea, metatarsal
I (56 mm), pedal ungual I, metatarsal II, metatarsal III (200 mm), metatarsal
IV, pedal phalanges, pedal unguals (Dong, Zhou and Zhang, 1983)
Comments- Dong et al. (1978) list Szechuanosaurus "yandonensis"
as a new species in a faunal list of taxa from the Wujiaba quarry of the Shangshaximiao
Formation. There is no description or illustration, making this a nomen nudum.
In 1983, Dong et al. note there was only a single large theropod skeleton in
the Wujiaba quarry, described by them as a neotype of Szechuanosaurus campi.
It can be implied that Dong et al. originally believed IVPP V237-239 to be a
new species of Szechuanosaurus, but later decided to include it in S.
campi.
Paul (1988) noted the teeth associated with CV 00214 lack roots, so were shed
by a scavenger. He placed the specimen in Metriacanthosaurus (along with
Yangchuanosaurus), as Metriacanthosaurus? sp.. He stated "there
are so many uncertainties about this beast that I balk at giving it a new name."
Yet in the earlier discussion of Metriacanthosaurus, Paul refers to the
species as M. carpenteri and states it belongs in a separate subgenus
than M. parkeri and M. shangyouensis. One may conclude Paul originally
intended to name the taxon, but later changed his mind and didn't catch all
the times he had used the name. In any case, "carpenteri" is a nomen
nudum since ICZN article 11.5 states "To be available, a name must be used
as valid for a taxon when proposed ..."
Chure (2000) used this specimen as the holotype of his new species "Szechuanoraptor
dongi" in his unpublished thesis. Names in theses aren't usually listed
in this website, and this one is only because it was later published by Glut
(2003). Glut's work includes a caveat to the effect that it is not available
to establish new taxonomy however, so the name remains unofficial. Chure referred
the specimens of Szechuanosaurus? zigongensis to "Szechuanoraptor
dongi" as well, but this seems incorrect, since they are from an earlier
formation and differ morphologically.
References- Dong, Chang, Li and Zhou, 1978. Note on a new carnosaur Yanchuangosaurus
shangyuanensis gen. et sp. nov.) from the Jurassic of Yangchuan District,
Szechuan Province. Ke Xue Tong Bao [Science Newsletter]. 23(5), 302-304.
Dong, Zhou and Zhang, 1983. Dinosaurs from the Jurassic of Sichuan Palaeontologica
Sinica Whole Number 162 New Series C, Number 23 Edited by Nanjing Institute
of Geology and Palaeontology and Institute of Vertebrate Paleontology and Paleoanthropology
Academia Sinica (pp. 1-136) Science Press Peking, 1983 43 plates.
Paul, 1988. Predatory Dinosaurs of the World. Simon & Schuster, New York.
Chure, 2000. A new species of Allosaurus from the Morrison Formation
of Dinosaur National Monument (UT-CO) and a revision of the theropod family
Allosauridae. PhD dissertation. Columbia University. 964 pp.
Rauhut, 2000. The interrelationships and evolution of basal theropods (Dinosauria,
Saurischia). Ph.D. dissertation, Univ. Bristol [U.K.], 1-440.
Glut, 2003. Dinosaurs: The Encyclopedia. Supplement 3.McFarland Press. 726 pp.
Szechuanosaurus? zigongensis
Gao, 1993
Bathonian-Callovian, Middle Jurassic
Xiashaximiao Formation, Sichuan, China
Holotype- (ZDM 9011) ten cervical vertebrae, cervical ribs, thirteen
dorsal vertebrae (series 1.17 m), dorsal ribs, five sacral vertebrae, twenty-five
caudal vertebrae, scapula, humerus (360 mm), radius (200 mm), ulna (240 mm),
distal carpal (46 mm), metacarpal I (62 mm), phalanx I-1 (92 mm), metacarpal
II (118 mm), phalanx II-1 (75 mm), metacarpal III (107 mm), metacarpal IV (52
mm), ilia (550 mm), pubes (580 mm), ischia (510 mm)
Paratypes- ?(ZDM 9012) maxilla, teeth
?(ZDM 9013) ten teeth
?(ZDM 9014) femur, tibia, fibula
Diagnosis- (after Rauhut, 2000) differs from Gasosaurus and Xuanhanosaurus
in the more rectangular deltopectoral crest and the proximal part of the humerus
being less expanded transversely; from Monolophosaurus and Eustreptospondylus
in the gradually sloping anterior rim of the maxilla and the lack of opisthocoelous
cervical vertebrae; from Piatnitzkysaurus in the gradually sloping anterior
rim of the maxilla and the less expanded proximal humerus; from Poekilopleuron
in the more strongly pronounced olecranon process of the ulna and the lack of
a medial process on the radius; from Metriacanthosaurus in the less steeply
sloping posterodorsal rim of the ilium; from Proceratosaurus in the more
massive and relatively shorter posterior part of the maxilla.
Comments- ZDM 9011 was discovered in 1984 and initially referred to Gasosaurus,
but described by Gao (1993) as a new species of Szechuanosaurus - S.
zigongensis. This was based on a number of characters, largely symplesiomorphies.
Chure (2000) referred these specimens to his new taxon "Szechuanoraptor
dongi" (based on CV 00214), needlessly creating a new species name, but
there appears to be little reason for such a referral. The specimens derive
from different formations and differ morphologically.
References- Gao, 1993. A new species of Szechuanosaurus from the
Middle Jurassic of Dashanpu, Zigong, Sichuan. Vertebrata PalAsiatica. 31(4):
308-314.
Chure, 2000. A new species of Allosaurus from the Morrison Formation
of Dinosaur National Monument (UT-CO) and a revision of the theropod family
Allosauridae. PhD dissertation. Columbia University. 964 pp.
Rauhut, 2000. The interrelationships and evolution of basal theropods (Dinosauria,
Saurischia). Ph.D. dissertation, Univ. Bristol [U.K.], 1-440.
Xuanhanosaurus Dong,
1984
X. qilixiaensis Dong, 1984
Bathonian-Callovian, Middle Jurassic
Xiashaximiao Formation, Sichuan, China
Holotype- (IVPP V6729) four anterior dorsal vertebrae, posterior dorsal
vertebra, dorsal neural arch, partial scapula, coracoid, humerus (265 mm), radius
(202 mm; proximal end lost), ulna (240 mm; lost), distal carpal I, distal carpal
II, distal carpal III, metacarpal I (52 mm), phalanx I-1 (84 mm), manual ungual
I (64 mm), metacarpal II (109 mm), phalanx II-1 (64 mm), metacarpal III (94
mm), phalanx III-1 (32 mm), phalanx III-2 (28 mm), metacarpal IV (50 mm)
Diagnosis- (from Rauhut, 2000) glenoid articular facet of humerus forms
a raised horizontal ridge that overhangs the humeral shaft posteriorly.
Comments- The supposed sternum is part of the coracoid (Rauhut, 2000).
The phylogenetic position is based on Holtz et al., (2004), though Rauhut (2000)
also found it to be a non-coelurosaur tetanurine.
References- Dong, 1984. A new theropod dinosaur from the Middle Jurassic
of Sichuan Basin. Vertebrata PalAsiatica. 22(3), 213-218.
Rauhut, 2000. The interrelationships and evolution of basal theropods (Dinosauria,
Saurischia). Ph.D. dissertation, Univ. Bristol [U.K.], 1-440.
Holtz, Molnar and Currie, 2004. Basal Tetanurae. In Weishampel, Dodson and Osmolska.
The Dinosauria Second Edition. University of California Press. 861 pp.
unnamed probable tetanurine (Russell, 1996)
Albian, Early Cretaceous
Gres Rouges Infracenomaniens, Morocco
Material- (CMN 41865) distal humerus (~385 mm, 156 mm wide)
Comments- Very similar to Xuanhanosaurus.
Reference- Russell, D.A. 1996. Isolated dinosaur bones from the Middle
Cretaceous of the Tafilalt, Morocco. Bulletin du Muse'um national d'Histoire
naturelle (4e se'r.) 18:349-402.