Scientific Publications of J.H. Epler
 (last updated 19 September 2014)
Please note that many of these publications are NOT available through me; these are marked with an *.  However, some of them may be ordered through their publishers or other sources; please follow the links!  Otherwise, drop me an email request.

1.   * Polk, J.L., J.H. Kennedy, W.A. Potter & J.H. Epler. 1976. The effects of flooding on populations of Chironomus Meigen (Diptera: Chironomidae) and Limnodrilus Claparedé (Oligochaeta: Tubificidae) in the Susquehanna River in the vicinity of Three Mile Island, Pennsylvania. Proc. Pa. Acad. Sci. 50:91-95.

2.   Epler, J.H. 1986. Book review: Chironomidae of the Holarctic region. Keys and Diagnoses. Vol. 1. Larvae; A catalogue of the chironomid genera and subgenera of the world including synonyms (Diptera: Chironomidae); A review of the genera Doithrix n. gen., Georthocladius Strenzke, Parachaetocladius Wülker and Pseudorthocladius Goetghebuer (Diptera: Chironomidae, Orthocladiinae). Proc. Ent. Soc. Wash. 88: 194-196.

3.   * Epler, J.H. 1986. Geographic distribution: Sphaerodactylus elegans. SSAR Herp. Rev. 17:27.

4.   Epler, J.H. 1986. A novel new Neotropical Nanocladius (Diptera: Chironomidae), symphoretic on Traverella (Ephemeroptera: Leptophlebiidae). Fla. Ent. 69:319-327. CLICK HERE to download a PDF version of this paper (a little over one Meg).

5.   Epler, J.H. 1986. Oukuriella, a new genus of Chironomidae (Diptera) from South America. Ent. Scand. 17:157-163.

6.   Epler, J.H. 1987. The larva of Radotanypus submarginella (Sublette) (Diptera:Chironomidae). Spixiana 9:285-287.

7.   * Epler, J.H. 1987. Revision of the Nearctic Dicrotendipes Kieffer, 1913 (Diptera: Chironomidae). Evol. Monogr. 9:102 pp + 37 plates.

8.   Epler, J.H. 1987. Notes on the Dicrotendipes (Diptera: Chironomidae) of Mexico, with descriptions of two new species. Ent. Scand. Suppl. 29:147-154.

9.   * Epler, J.H. 1988. Biosystematics of the genus Dicrotendipes Kieffer, 1913 (Diptera: Chironomidae: Chironominae) of the World. Mem. Am. Ent. Soc. 36:1-214.

10.  Epler, J.H. 1988. A reconsideration of the genus Apedilum Townes, 1945 (Diptera: Chironomidae). Spixiana Suppl. 14:105-116.  For a scanned PDF version (940 KB) of this paper, click here.

11.  * Epler, J.H. 1990. A quick guide to chironomid identification literature. Bull. N. Am. Benth. Soc. 7:238-246.

12.  * Livingston, R.J., J.H. Epler, F. Jordan, Jr., W.R. Karsteter, C.C. Koenig, A.K.S.K. Prasad & G. Ray. 1990. Ecology of the Choctawhatchee River System. Pp. 247-274 in Livingston, R.J. (ed.). The Rivers of Florida. Springer-Verlag, New York,.

13.  * Epler, J.H. 1992. Identification Manual for the Larval Chironomidae (Diptera) of Florida. FL Dept. Environ. Reg., Orlando, FL. 302 pp.

14.  Epler, J.H. & L.C. Ferrington, Jr. 1994. The immature stages of Paratendipes basidens Townes (Diptera: Chironomidae: Chironominae). J. Kans. Ent. Soc. 67: 311-317.

15.  Mason, W.T., Jr., R.A. Mattson & J.H. Epler. 1994. Benthic macroinvertebrates and allied macrofauna in the Suwannee River and estuary ecosystem, Florida. Florida Scientist 57: 141-160.

16.  * Epler, J.H. 1995. Identification Manual for the Larval Chironomidae (Diptera) of Florida. Revised edition. FL Dept. Environ. Protection, Tallahassee, FL. 317 pp.

17.  * Epler, J.H. 1995. Book review: The Chironomidae. The biology and ecology of non-biting midges. J. N. Am. Benthol. Soc. 14: 461-462.

18.  * Epler, J.H. 1995. Book review: Chironomids - from genes to ecosystems. J.N. Am. Benthol. Soc. 14: 674-675.

19.  Epler, J.H. & C.L. de la Rosa. 1995. Tempisquitoneura, a new genus of Neotropical Orthocladiinae (Diptera: Chironomidae) symphoretic on Corydalus (Megaloptera: Corydalidae). J. N. Am. Benthol. Soc. 14: 50-60.

20.  Hubbard, M.D., M.L. Pescador, A.K. Rasmussen, I.S. Askevold, J. Jones, R.W. Flowers & J.H. Epler. 1995. The freshwater macroinvertebrates of Florida: a guide to references for their identification. Fl. Ent. 78: 161-177.  NOTE: a PDF version of this paper can be found on-line at

21.  * Mason, W.T., Jr. & J.H. Epler 1995. Guide to taxonomic literature on benthic invertebrates of the southeastern United States. Ecology Support Publ. ES 95-02, 36 pp.

22.  * Mattson, R.A., J.H. Epler & M.K. Hein. 1995. Description of benthic communities in karst, spring-fed streams of north central Florida. J. Kans. Ent. Soc. 68 suppl: 18-41.

23.  Epler, J.H. 1996. New species of Oukuriella Epler (Diptera: Chironomidae) from Costa Rica. Hydrobiologia 318: 3-11.

24.  Epler, J.H. 1996. A new species of Dicrotendipes Kieffer (Diptera: Chironomidae) from Costa Rica. Hydrobiologia 318: 13-15.

25.  * Epler, J.H. 1996c. Identification Manual for the Water Beetles of Florida (Coleoptera: Dryopidae, Dytiscidae, Elmidae, Gyrinidae, Haliplidae, Hydraenidae, Hydrophilidae, Noteridae, Psephenidae, Ptilodactylidae, Scirtidae). FL Dept. Environ. Protection, Tallahassee, FL. 257 pp.

26.  Epler, J.H., A.D. Harrison & L. Hare. 1999. Acinoretracus, a new Afrotropical genus for some species previously placed in Dicrotendipes (Diptera: Chironomidae: Chironominae). Tijd. voor Ent. 141: 209-220.

27.  * Epler, J.H. 1999.  An Introduction to the Taxonomy and Identification of Larval Chironomidae. Workbook for the 1999 North American Benthological Society Technical Workshop on Larval Midge Taxonomy, Duluth, MN, 25 May 1999.  54 pp. Available through NABS; click here for ordering information.

28.  Epler, J.H. & W.J. Janetzky. 1999. A new species of Monopelopia (Diptera: Chironomidae) from phytotelmata in Jamaica, with preliminary ecological notes.  J. Kans. Ent. Soc.  71: 216-225. PDF download (371 KB)

29.  Epler, J.H., J.P. Cuda & T.D. Center.  2000.  Redescription of Cricotopus lebetis (Diptera: Chironomidae), a potential biocontrol agent of the aquatic weed hydrilla (Hydrocharitaceae).  Fl. Ent. 83: 171-180.  A PDF version of this paper (273Kb) may be obtained by clicking here.

30.  Epler, J.H. 2001. Identification Manual for the larval Chironomidae (Diptera) of North and South Carolina.  A guide to the taxonomy of the midges of the southeastern United States, including Florida. Special Publication SJ2001-SP13.  North Carolina Department of Environment and Natural Resources, Raleigh, NC, and St. Johns River Water Management District, Palatka, FL. 526 pp.  A PDF version can be downloaded by clicking here.

31.  Lester, G.T., B.J. Krestian & J.H. Epler. 2004 [2003]. First Nearctic records of Tempisquitoneura (Diptera: Chironomidae: Orthocladiinae) from Arizona, USA. Ent. News 114: 117-119.

32. Wang, X., Z. Liu & J.H. Epler.  2004.  New species of Bryophaenocladius Thienemann from the Nearctic Region (Diptera: Chironomidae: Orthocladiinae).  Zootaxa 581: 1-10. 

33.  Barman, E.H. & J.H. Epler. 2005. Errors in descriptions of larvae of Thermonectus Dejean (Coleoptera: Dytiscidae). Southeastern Naturalist 4: 119-120.

34. Epler, J.H., D.H. Ray & T.A. Thom. 2005. New distribution records for water beetles (Coleoptera: Elmidae, Gyrinidae) in Florida. Col. Bull. 59: 270-271. 

35. Paller, M.H., F.D. Martin, L.D. Wike & J.H. Epler. 2005. Biological assessment of slope wetlands on the Coastal Plain of South Carolina. J. Freshwater Ecol. 20: 247-262.

36. Epler, J.H. 2006. Identification Manual for the Aquatic and Semi-aquatic Heteroptera of Florida (Belostomatidae, Corixidae, Gelastocoridae, Gerridae, Hebridae, Hydrometridae, Mesoveliidae, Naucoridae, Nepidae, Notonectidae, Ochteridae, Pleidae, Saldidae, Veliidae). Florida Department of Environmental Protection, Tallahassee, FL. 186 pp.  Click here to download from FDEP server.

37. Hribar, L.J. & J.H. Epler. 2007. Collection records for some Chironomidae (Diptera) in the Florida Keys. Fl. Scientist 70: 110-112.

38. Schärer, M.T. & J.H. Epler.  2007.  Long-range dispersal possibilities via sea turtle: a case for Clunio and Pontomyia (Diptera: Chironomidae) in Puerto Rico.  Ent. News 118 (3): 273-277.

39. Donato, M., A.C. Paggi & J.H. Epler. 2008. New record, geographic variation, and redescription of Apedilum elachistus Townes (Diptera: Chironomidae: Chironominae). Studies on Neotropical Fauna and Environment: 43: 125-134. 

40. Hribar, L.J., J.H. Epler, J. Martin & J.E. Sublette.  2008.  Chironomus columbiensis (Diptera: Chironomidae) new to the fauna of the United States. Fl. Ent. 91: 470-471.

41. Epler, J.H. 2009. More new distribution records for Florida water beetles (Coleoptera: Dytiscidae, Elmidae, Hydrophilidae, Scirtidae), with additional notes on Scirtes oblongus Guérin-Méneville. Insecta Mundi 0087: 1-4.

42. Spies, M., T. Andersen, J.H. Epler & C.N. Watson, Jr. 2009. Chironomidae (non-biting midges). Pp. 437-480 in Brown, B.V. et al.  Manual of Central American Diptera: Volume 1. NRC Research Press, Ottawa, Ontario, Canada. 714 pp.

43.  Thompson, F.C. & J.H. Epler.  2009 (2008).  Diptera (Flies) of Attu: The first assessment of the entomofauna of the last place on earth.  Studia dipterologica 15: 267-275.

44.  Krestian, B.J, E. Kosnicki, P.H. Spindler, S. Stringer & J.H. Epler. 2010. First Nearctic records of Oliveiriella Wiedenbrug and Fittkau, with new distributional records for two other New World species of Orthocladiinae (Diptera: Chironomidae).  Ent. News 120: 349-362.

45.  Epler, J.H.  2010.  The Water Beetles of Florida - an identification manual for the families Chrysomelidae, Curculionidae, Dryopidae, Dytiscidae, Elmidae, Gyrinidae, Haliplidae, Helophoridae, Hydraenidae, Hydrochidae, Hydrophilidae, Noteridae, Psephenidae, Ptilodactylidae and Scirtidae.  Florida Department of Environmental Protection, Tallahassee, FL. 399 + iv pp.  Click here to download from FDEP server.

46.  Pfrender, M.E, C.P. Hawkins, M. Bagley, G. W. Courtney, B. R. Creutzburg, J.H. Epler, S. Fend, L.C. Ferrington, Jr., P.L. Hartzell, S. Jackson, D.P. Larsen, C.A. Levesque, J.C. Morse, M. Petersen, D. Ruiter, D. Schindel & M. Whiting. 2010. Assessing macroinvertebrate biodiversity in freshwater ecosystems: advances and challenges in DNA-based approaches. Quarterly Review of Biology 85: 319-340.

47.  Epler, J.H. 2010. Phytotelmatocladius,  a new genus from bromeliads in Florida and Brazil (Diptera: Chironomidae: Orthocladiinae). Pp. 285-293 in Ferrington, L.C., Jr. (ed.) Proceedings of the XV International Symposium on Chironomidae.  Chironomidae Research Group, University of Minnesota, St. Paul, MN. 385 + vii pp. 

48.  Epler, J.H.  2011.  Now see-um.  Florida Wildlife. January/February, pp. 30-31.

49. Copeland, R.S., E. Nkubaye, B. Nzigidahera, J.H. Epler, J.P. Cuda & W.A. Overholt. 2012.   The diversity of Chironomidae (Diptera) associated with Hydrilla verticillata (Alismatales: Hydrocharitaceae) and other aquatic macrophytes in Lake Tanganyika, Burundi. Annals of the Entomological Society of America 105: 206-224.

50.  Epler, J.H. 2012.  A brachypterous Bryophaenocladius (Diptera: Chironomidae: Orthocladiinae) with hypopygium inversum from Heggie’s Rock, Georgia, U.S.A.  Zootaxa 3355: 51-61.

51.  Gray, E.W., C. Royals, J.H. Epler, R.D. Wyatt, B. Brewer & R. Noblet.  2012.  Chironomus calligraphus (Diptera: Chironomidae), a new pest species in Georgia. Journal of the American Mosquito Control Association 28: 258-259.

52.  Copeland, R.S., B. Gidudu, F. Wanda, J.H. Epler, J.P. Cuda & W.A. Overholt.  2012.  Chironomidae (Insecta: Diptera) collected from Hydrilla verticillata (Hydrocharitaceae) and other submersed aquatic macrophytes in Lake Bisina and other Ugandan lakes.  Journal of East African Natural History 101: 29-66.

53.  Epler, J.H.  2012.  A new species of Scirtes (Coleoptera: Scirtidae) from southern Florida and the Caribbean.  Zootaxa 3530: 77-82.

54.  Megna, Y.S. & J.H. Epler.  2012.  A review of Copelatus from Cuba, with the description of two new species (Coleoptera: Dytiscidae: Copelatinae). Acta Entomologica Musei Nationalis Pragae 52: 383-410.

55.  Andersen, T., Cranston, P. S. & Epler, J. H. (Sci. eds).  2013. The larvae of Chironomidae (Diptera) of the Holarctic Region — Keys and diagnoses. Insect Systematics & Evolution, Supplement 66: 1- 571.

56.  Cranston, P. S., & Epler, J.H. 2013. 5. The larvae of Tanypodinae (Diptera: Chironomidae) of the Holarctic region — Keys and diagnoses. pp. 39-136.  In: Andersen, T., Cranston, P. S. & Epler, J. H. (Sci. eds): The larvae of Chironomidae (Diptera) of the Holarctic Region — Keys and diagnoses. Insect Systematics & Evolution, Supplement 66: 1- 571.

57.  Andersen, T., Sæther, O. A., Cranston, P. S. & Epler, J. H. 9. The larvae of Orthocladiinae (Diptera: Chironomidae) of the Holarctic Region — Keys and diagnoses. pp. 189–386. In: Andersen, T., Cranston, P. S. & Epler, J. H. (Sci. eds): The larvae of  Holarctic Chironomidae (Diptera) of the Holarctic Region — Keys and diagnoses. Insect Systematics & Evolution, Supplement 66: 1- 571.

58.  Epler, J.H., Ekrem, T. & Cranston, P.S. 2013. 10. The larvae of Chironominae of the Holarctic Region — Keys and diagnoses. pp. 387-556. In: Andersen, T., Cranston, P. S. & Epler, J. H. (Sci. eds): The larvae of Chironomidae (Diptera) of the Holarctic Region — Keys and diagnoses. Insect Systematics & Evolution, Supplement 66: 1- 571.

59.  Epler, J.H. 2014. Identification Guide to the Larvae of the Tribe Tanytarsini (Diptera: Chironomidae) in Florida.  Florida Department of Environmental Protection, Tallahassee, FL. 77 + iv pp.  Click here to download from FDEP server.

60.  Oh, J.T., Epler, J.H. & Bentivegna, C.S.  2014.  A rapid method of species identification of wild chironomids (Diptera: Chironomidae) via electrophoresis of hemoglobin proteins in sodium dodecyl sulfate polyacrylamide gel (SDS–PAGE).  Bulletin of Entomological Research 104: 13 pp.  available on CJO2014. doi:10.1017/S0007485314000431.

61.  Milensky, C.M, Austin, C.J., Epler, J.H., Gebhard, C.A. & Graves, G.R.  2014.  Chironomid midge hatch leads to mass mortality event for Chimney Swifts (Chaetura pelagica).  Banisteria 43: 99-101.

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